Honey Bee Biology
Before you get bees, it helps to really understand how they work. A solid grasp of bee biology makes everything else about beekeeping easier. You don’t need to be a scientist, but knowing what’s going on inside the hive helps you make better decisions and avoid a lot of mistakes.
Three Types of Bees in a Hive
A honey bee colony is a superorganism, meaning it functions like one big living thing. It’s made up of three types of bees (called castes):
Queen Bee : She lays all the eggs. One queen per hive.
Worker Bees : All female, infertile, and make up 90%+ of the hive. They do everything else: clean, defend, feed, and forage.
Drones : The male bees. Their only job is to mate with a queen.
Each of these bees has a different role, body type, and lifespan. They work together as a team. No bee can survive very long on its own.
The Basics of Bee Anatomy
All honey bees have three main body parts: head, thorax, and abdomen.
Head : Has five eyes, two antennae, and a tongue (proboscis). The antennae help bees smell, taste, feel, and even detect temperature and humidity.
Thorax : Where the wings and legs attach. This part powers flight.
Abdomen : Contains the stomach and stinger (for queens and workers).
Worker bees have a barbed stinger, so when they sting, it usually means death for them. Queen stingers are smooth and can be used more than once, but they’re only used on other queens.
Worker Bee Life
Worker bees cycle through jobs as they age. Early on, they clean cells and feed larvae. Later, they take on guarding duties, and eventually become foragers. In spring and summer, they live about 6 weeks. In winter, when they don’t fly much, they can live several months.
Drones
Drones don’t forage or defend the hive. They mate with queens from other hives and die right after mating. During a nectar dearth or in winter, they get kicked out of the hive since they don’t contribute and just use up resources.
Queen Bee
The queen’s job is to lay eggs—up to 2,500 per day at her peak. She lives for 1-3 years, depending on how well-mated she is. If she starts to run out of sperm or fails, the colony will replace her. Worker bees rear new queens from fertilized eggs by feeding them only royal jelly.
How Bees Are Born
Bees go through four stages: egg, larva, pupa, and adult. The timeline depends on the caste:
Worker : 21 days
Queen : 16 days
Drone : 24 days
The queen lays an egg in a wax cell. After hatching, the larva is fed royal jelly and later bee bread (a mix of pollen and nectar). It spins a cocoon, pupates, and finally emerges as an adult.
Whether a fertilized egg becomes a queen or a worker depends on what it's fed and the size of the cell it's laid in. Bigger cells mean drones, and smaller ones mean workers.
What a Colony Needs to Thrive
Honey bee colonies rely on four resources:
Pollen : Protein source used to feed young bees. Stored as bee bread.
Nectar : Collected and converted into honey. It’s their energy source.
Water : Used to cool the hive and dilute honey for feeding.
Propolis : Made from plant resins. Used to seal cracks, sanitize the hive, and even mummify invaders.
Swarming: Colony Reproduction
When the hive gets crowded or conditions are right, bees prepare to swarm. Half the colony leaves with the old queen to start a new colony. The bees left behind raise a new queen. It’s how colonies reproduce.
Bee Communication
Bees communicate mostly through pheromones and movement. Their antennae are packed with sensors. They also do the famous waggle dance to tell others where to find food.
Key pheromones include:
Queen pheromone : Keeps the hive orderly and tells workers a queen is present.
Alarm pheromone : Smells like bananas. Warns of threats.
Nasanov pheromone : Helps bees find their way back to the hive.
Footprint pheromone : Helps mark comb and discourage new queen cells.
Brood pheromone : Comes from larvae. Tells workers to feed them and affects foraging behavior.
If queen or brood pheromones go missing too long, some workers may develop ovaries and start laying unfertilized eggs—which only turn into drones. This is called a laying worker colony, and it’s tough to reverse.
This chapter sets the foundation. Next up: where bees live and how they build their comb.
A Honey Bee’s Home
When we talk about a honey bee’s home, we’re not just talking about the wooden hive you set up. That’s just the structure. The real home is the comb the bees build inside. Beeswax comb is where everything happens—brood rearing, food storage, and communication.
What is Beeswax Comb?
Beeswax is produced by worker bees aged 12–18 days. They convert honey into wax using glands on the underside of their abdomen. It comes out in thin flakes, which they chew and form into hexagon-shaped cells.
To make just one pound of wax, bees need to eat about eight pounds of honey. That’s why comb-building only happens during strong nectar flows. If natural nectar isn’t available, you can feed sugar syrup to help them build.
What Bees Use the Comb For
Brood nest : Where eggs are laid and baby bees grow.
Food storage : Pollen and nectar are stored here. Pollen gets turned into bee bread. Nectar gets turned into honey.
Cooling system : Bees fan water in the comb to cool the hive in summer.
Communication : Bees use vibration through the comb to send messages.
Comb Color Changes
Fresh comb starts out white, almost see-through. It gets darker over time from use—especially from pollen oils, propolis lining, and leftover cocoons from brood cycles. Older comb can turn brown or black. That’s normal, but many beekeepers rotate out older frames yearly to keep things clean and healthy.
How Bees Build in the Wild
In nature, bees look for enclosed, sheltered spaces with small entrances—tree trunks, owl boxes, sheds, even wall voids. The entrance size helps them defend against pests.
They build the comb in vertical sheets, creating a central brood nest surrounded by food stores. Drones are usually raised around the outer edges of the brood nest. If the colony gets cold, bees cluster in the middle to protect the queen and worker brood first. Drone brood on the outer edge acts as a buffer.
Understanding Bee Space
Bees follow strict spacing rules. If the space is too small (< 1/4"), they fill it with propolis. If it’s too big (> 3/8"), they build comb. This concept, called “bee space,” is the reason modern hives work so well. It lets you inspect the hive without destroying comb.
Identifying What You’re Seeing
Here’s how to tell what’s what when you open your hive:
Comb : Starts white, darkens with use.
Queen : Long abdomen, moves with purpose. Wings shorter than abdomen.
Drone : Large, rounded body. Big eyes meet at the top of the head.
Worker : Smaller than the others. Hairy bodies and tattered wings as they age.
Eggs : Tiny, like a grain of rice, stuck to the bottom of cells.
Larvae : Look like tiny white grubs curled in cells.
Pupae : Capped brood. Worker caps are flat; drone caps are dome-shaped.
Bee Bread : Shiny, packed pollen with layers of color.
Honey : Sealed with wax. Doesn’t spill out when shaken.
Nectar : Glossy and wet. Shakes out easily.
Queen cells look like peanuts and stick out from the comb. If they’re uncapped and empty, they’re called nubbins—practice cells the bees might use later.
Here’s Chapter 3 for your reference:
Honey Bee Nutrition
To be a good beekeeper, you have to understand what bees eat and when they need help. Nutrition impacts everything: brood health, colony strength, and honey production.
What Bees Need
Honey bees need macronutrients (carbohydrates, proteins, lipids) and micronutrients (vitamins and minerals) just like we do.
Carbohydrates give them energy. Bees get it from nectar and honey.
Protein builds and repairs bodies. They get it from pollen.
Lipids (fats) support brood development and healthy cell function. Found in pollen.
Vitamins & minerals come from pollen, some nectar, and water. Darker honeys usually have more minerals.
Who Needs What
Larvae need lots of protein and royal jelly.
Worker bees need energy to fly and protein to develop glands.
Queens get a steady diet of royal jelly.
Drones don’t gather food, but they consume a lot of it.
Poor nutrition during development weakens bees for life. If a weak generation emerges during honey flow, you can end up with no surplus and a hungry hive going into winter.
Where Bees Get Their Food
Bees forage up to 3–5 miles to gather:
Pollen (protein and fat)
Nectar (carbs)
Water (to cool the hive and dilute honey)
Plant resins (to make propolis)
They use scent and UV vision to locate flowers. Many flowers have UV markings (nectar guides) visible only to bees. This directs them to nectar sources efficiently.
Waggle Dancing for Dinner
Once a bee finds a good food source, she comes back to the hive and performs a waggle dance. The direction, duration, and angle of the waggle tells other bees where to go. It's a GPS system in the dark.
Bees also mark good food spots and the hive entrance with pheromones so others can find their way.
Here’s Chapter 4: Getting Started with Beekeeping in full:
Getting Started with Beekeeping
Starting with bees doesn’t have to be overwhelming, but it helps to go in with a plan. Here's a rundown of what you need, what to expect, and how to set yourself up for success.
Choosing Your Hive Equipment
The most common setup is the Langstroth hive —stacked boxes with removable frames. It’s easy to manage and inspect.
Basic components you’ll need:
Bottom board (solid or screened)
Entrance reducer (helps control airflow and pests)
Deep boxes for brood (usually 1–2)
Medium or shallow boxes for honey
Frames with foundation (waxed plastic or wired wax)
Inner cover and outer cover
Other hive types like top-bar or horizontal hives can work too, but parts and help may be harder to find.
Protective Gear
Don’t skimp here. A good veil or suit helps you relax during inspections. Gloves are great when you’re starting out.
Essentials:
Bee suit or jacket with veil
Gloves (optional but recommended)
Closed-toe shoes or boots
Basic Tools
Just a few essentials will do:
Hive tool (your must-have for every visit)
Smoker (helps calm the bees)
Bee brush (for gently moving bees)
Choosing a Location
Good hive placement matters:
Morning sun exposure
Wind protection
Good drainage (avoid low, wet spots)
Nearby water source (shallow dish with rocks works great)
Room behind the hive to stand and inspect
Avoid placing hives near doors, patios, or busy walkways.
Buying Bees
You can start with:
Nucs (nucleus colonies): Come with bees, a queen, brood, and food—ideal for beginners.
Packages : A box of bees with a caged queen. More setup needed, but widely available.
Swarms : Free if you catch them, but less predictable.
When to Start
Install your bees in early spring when daily temps are consistently above 50°F. That gives them a full season to grow before winter.
Take a Class or Find a Mentor
You’ll learn way more, way faster. Join a local bee club, take a course, or ask another beekeeper for help. Watching someone open a hive makes everything less intimidating.
Let me know if you want the next chapter too.
Here’s Chapter 5: Hive Inspections :
Hive Inspections
Once your bees are installed and flying, you’ve got to check in on them. Regular hive inspections help you keep your colony healthy, prevent swarming, and catch issues before they get out of control. But don’t worry—this doesn’t mean opening the hive every day.
When to Inspect
Check your hive every 7–14 days during spring and summer. That’s often enough to track brood patterns and spot early signs of swarming. In fall, inspect every few weeks. In winter, just observe flight and check food stores without opening the hive.
Avoid inspections when:
It’s rainy or windy
It’s below 60°F
The nectar flow has stopped and bees are cranky
Mid-morning to early afternoon on warm, sunny days is ideal. Foragers will be out working, so fewer bees will be home.
What You’re Looking For
Each inspection should be quick and focused. Here’s a checklist:
Is the queen present? (You don’t have to see her—just look for eggs.)
Is there a healthy brood pattern? (Solid, even patches of capped brood)
Any signs of swarming? (Queen cells, crowded frames)
Are they storing honey and pollen?
Any pests or diseases? (Mites, beetles, wax moths)
It gets easier to read a hive the more you do it. Trust your gut—if something seems off, it probably is.
Tips for Smooth Inspections
Use your smoker! A few gentle puffs at the entrance and under the lid go a long way.
Work slowly and confidently. Jumpy movements rile up bees.
Start with the outer frames and work inward.
Don’t crush bees—gently move frames and lids.
Stay calm. If bees get aggressive, close up and try another day.
Taking Notes
Keep a notebook, app, or printed checklist handy. Log what you saw: brood pattern, number of frames covered in bees, queen sighting, any concerns. These notes will help you see trends and plan treatments or feeding.
Let me know if you want Chapter 6 next.
Here’s Chapter 6: Common Problems in the Hive :
Common Problems in the Hive
Even strong hives can run into trouble. The key is learning how to spot problems early and knowing how to respond.
Varroa Mites
These external parasites attach to adult bees and developing brood. They weaken bees and spread viruses. Left untreated, they’ll collapse the hive.
What to look for:
Mites on drone brood or adult bees
Deformed wings or stunted bees
Spotty brood pattern
What to do:
Test mite levels regularly with sugar rolls or alcohol washes
Treat with approved methods (like Apivar, oxalic acid, or formic acid) when thresholds are met
Small Hive Beetles
These beetles lay eggs in the hive. Their larvae ruin comb and ferment honey.
What to look for:
Slime or a fermented smell
Larvae crawling on comb
Beetles running across frames
What to do:
Use beetle traps
Keep hives strong and in full sun
Don’t leave empty equipment lying around
Wax Moths
These pests move in when a hive is weak or dead. They destroy comb, chew wood, and leave silky tunnels and webbing.
What to do:
Keep hives strong
Freeze unused frames before storage
Queen Issues
Queens can die, fail, or be poorly mated.
What to look for:
No eggs or only drone brood
Multiple eggs per cell (a sign of laying workers)
Spotty or missing brood
What to do:
Combine with another hive
Requeen with a mated queen
Add a frame of eggs from a good hive to encourage them to raise a new queen
Swarming
A natural process, but it weakens your hive.
What to look for:
Queen cups or queen cells
Nectar being stored in the brood nest
What to do:
Make a split
Reorganize frames to give space
Remove excess queen cells (if appropriate)
Robbing
Other bees will steal honey when there’s little nectar available.
What to look for:
Frenzied activity at the entrance
Bees fighting
Torn cappings inside the hive
What to do:
Reduce entrance size
Don’t leave syrup or wet supers out
Use robbing screens if needed
Let me know if you’d like Chapter 7.
Here’s Chapter 7: Feeding Your Bees :
Feeding Your Bees
Bees prefer to feed themselves from natural forage, but there are times when you’ll need to step in and give them a hand. Feeding helps build up young colonies, sustain hives during nectar dearths, and support bees during winter prep.
When to Feed
Feed your bees when:
You install a new package or nuc
Nectar sources are low (especially in late summer or fall)
You’re preparing bees for winter and they’re light on stores
You’re trying to stimulate brood production in spring
Don’t feed when honey supers are on —you don’t want sugar syrup mixed into your harvest.
What to Feed
1:1 Sugar Syrup (1 part sugar to 1 part water)
Use in spring to stimulate brood rearing.
2:1 Sugar Syrup (2 parts sugar to 1 part water)
Use in fall to help bees store up for winter.
Dry Sugar or Candy Boards
Used in winter when it’s too cold for syrup.
Pollen Substitute or Supplement
Feed when natural pollen is scarce (like early spring or droughts).
How to Feed
Entrance Feeders
Simple but can attract robbers—best used when it’s warm and there’s no dearth.
Top Feeders (Bucket or Hive-top)
Great for larger volumes. Reduces robbing risk.
Frame Feeders
Go inside the brood box. Effective, but bees can drown if it’s not a good design.
Candy Boards or Dry Sugar
Used in winter directly over the cluster.
Feeding Tips
Always use white granulated sugar (never brown sugar, honey, or molasses)
Keep feeders clean to avoid mold or fermentation
Only feed when needed—don’t make bees reliant on it
Want me to keep going with Chapter 8?
Here’s Chapter 8: Beekeeping Through the Seasons :
Beekeeping Through the Seasons
Beekeeping isn’t a set-it-and-forget-it hobby. What you do depends on the season. Bees have different needs throughout the year, and your job is to keep up with them as the seasons change.
Spring
Spring is all about growth. Colonies expand rapidly and need space.
Watch for swarming—colonies build up fast
Add boxes as the brood nest fills
Feed 1:1 syrup if nectar is low
Requeen weak hives
Monitor mites and treat if needed
Make splits if hives are booming
Summer
This is often the main honey flow—if you’re lucky.
Add honey supers before they fill up
Watch for congestion and swarming signs
Provide water and ventilation
Watch for robbing and queen issues
Keep an eye on mites—mid-summer spikes are common
Fall
Fall is about preparing for winter. You’re helping bees raise a strong final batch of workers.
Feed 2:1 syrup if stores are low
Optionally feed pollen substitute to boost brood
Remove extra boxes
Reduce entrances and install mouse guards
Finish mite treatments early before temps drop
Winter
In winter, bees cluster for warmth. There’s not much to do—but don’t disappear entirely.
Make sure hives are ventilated and protected from wind
Check weight by lifting the back of the hive
Add candy boards or dry sugar if needed
Watch for moisture buildup
Avoid opening the hive unless absolutely necessary
Want to see Chapter 9 next?
Here’s Chapter 9: Splitting Hives and Expanding Your Apiary :
Splitting Hives and Expanding Your Apiary
Once you’re comfortable with one or two hives, you might want more. Expanding your apiary can be done without buying more bees—just split the colonies you already have. Splitting also helps prevent swarming and gives you insurance against queen failure.
Why Split a Hive?
Prevent swarms in strong colonies
Raise new queens
Create backup hives in case one fails
Grow your operation without buying more bees
When to Split
Late spring to early summer is ideal. You want:
Warm weather
Drones in the air (for queen mating)
A strong hive with lots of brood and bees
How to Make a Simple Split (Walk-Away Method)
Find a strong hive with eggs, brood, and plenty of bees
Move 3–5 frames into a new hive box (include brood, eggs, honey, and pollen)
Shake in some extra bees if needed
Close up both hives
Walk away—the new colony will raise its own queen
Or you can introduce a mated queen instead for quicker results.
What You’ll Need
Extra hive body and lid
Bottom board
Feeders (if there's no nectar flow)
Queen excluder (optional for vertical splits)
Aftercare
Check in 7–10 days for queen cells
Don’t disturb capped queen cells
Give them 3–4 weeks to raise a queen, let her mate, and start laying
Be patient. Rushing it can disrupt the queen's development and lead to failure.
Let me know if you'd like Chapter 10 next.
Here’s Chapter 10: Collecting and Catching Swarms :
Collecting and Catching Swarms
Catching a swarm is one of the most exciting parts of beekeeping. It’s a chance to get free bees, usually with a mated queen, and it’s often a sign of a healthy, productive colony.
Why Catch a Swarm?
Free bees
Usually healthy and ready to build
Already have a queen
Fun and rewarding experience
When Do Bees Swarm?
Most swarming happens in spring and early summer, especially after a few warm days following rain. Peak hours are mid-morning to early afternoon on calm, sunny days.
What You Need
Ventilated box or nuc
Pruners or loppers
Bee brush or feather
Spray bottle with sugar water
Ladder (if they’re up high)
Protective gear (recommended)
Optional but helpful:
Queen clip or cage
Swarm lure (lemongrass oil works well)
Tarp or sheet to catch falling bees
How to Catch a Swarm
Find the cluster, usually hanging from a branch or structure
Shake or brush the bees gently into your container
Watch for fanning behavior—that means the queen’s inside
Close the box and take them to your apiary
If they fly off, the queen may still be outside. Give it another shot.
After the Catch
Install them into a hive within 24 hours
Feed 1:1 sugar syrup if no nectar is flowing
Check in a few days for eggs or brood
Tips
Stay calm—swarms are usually very gentle
Don’t chase them mid-flight
Tell local pest control or neighbors you’re available for swarm calls
Want to keep going with Chapter 11?
Harvesting Honey
Harvesting honey is one of the biggest rewards in beekeeping—but timing, technique, and care for your bees all matter. You want to do it right, both for your bees and your jars.
When to Harvest
Only harvest fully capped honey —that’s honey sealed with white wax. If it’s still shiny and uncapped, it has too much moisture and can ferment.
Most folks harvest:
Late spring and/or late summer
After supers are full and nectar flow slows down
How Much to Take
Leave at least 60 pounds of honey for your bees going into winter (about one full deep box). Don’t take more than they can spare.
How to Harvest
-
Clear the bees from the honey super:
Use a fume board
Try a bee escape board
Gently blow them off or brush them
Uncap the frames with a hot knife, cappings scratcher, or uncapping roller
Extract using a centrifugal extractor (borrow one if needed)
Strain the honey through a filter or cheesecloth
Bottle in clean, dry containers
What If You Don’t Have an Extractor?
You can crush and strain comb:
Cut the comb from the frame
Mash it in a strainer
Let it drip through a filter into a bucket
Takes longer, but works fine for small batches.
Cleaning Up
Return sticky frames to the bees—they’ll clean them up
Don’t leave them out in the open or you’ll trigger robbing
Final Tips
Label jars with harvest date and location
Store in a cool, dry place
If it crystallizes, warm gently in hot water (not the microwave)
Never feed harvested honey to other colonies—it could spread disease
Let me know if you want Chapter 12 next.
Here’s Chapter 12: When a Hive Dies (Dealing with Deadouts) :
When a Hive Dies (Dealing with Deadouts)
Losing a hive is frustrating—but it happens, even to experienced beekeepers. The key is to learn from it and use the loss to make your other hives stronger.
Common Reasons Hives Die
Starvation – They ran out of food in late winter or early spring
Varroa mites – Weakened the bees or spread viruses
Queen failure – The colony became queenless and didn’t recover
Moisture – Condensation dripped onto the cluster and chilled them
Too small a cluster – Not enough bees to stay warm through winter
Signs of a Deadout
No live bees in the hive
Bees headfirst in cells (starvation)
Moldy or chewed comb
Wax moth or mouse damage
Spotty brood patterns before death
Dead bees at the entrance or on the bottom board
What to Do With the Equipment
Clean and reuse : If no sign of disease
Freeze the frames : Kills pests like wax moths or beetles
Scorch woodenware if there’s mold
Discard if you suspect American Foulbrood (look for ropey brood)
Reusing Honey
If it’s clean and unfermented, you can feed it back to other colonies or extract it for yourself. Never feed questionable frames to bees.
Learn From It
Ask yourself:
Was the hive light or heavy?
Did you check mite loads in fall?
How was the queen performance?
Every deadout gives you info. Don’t take it personally—use it to improve.
Want Chapter 13 next?
Here’s Chapter 13: Final Gear + Tips Roundup :
Final Gear + Tips Roundup
As you wrap up your first year (or start planning for it), here’s a quick list of tools, gear, and habits that make beekeeping smoother, safer, and more enjoyable.
Must-Have Gear
Hive tool – your #1 essential
Smoker – calms bees and buys you time
Bee suit or jacket with veil – for comfort and confidence
Gloves – especially early on
Feeder – for spring and fall support
Spare boxes and frames – always handy
Notebook or app – to log what you see
Handy Extras
Frame holder – keeps frames safe while inspecting
Queen marking pen or cage – easier to track your queen
Mite test kits – for sugar roll or alcohol wash
Tarp or old sheet – for swarm catching or messy jobs
Bucket with lid – for hauling tools or wax scraps
Good Habits
Test for mites regularly—not just when bees look sick
Keep inspection notes so you can see patterns
Start small and grow slow
Ask for help—it’s part of beekeeping
Watch your bees daily, even without opening the hive
Where to Learn More
Join a local bee club
Take a beginner class
Follow trustworthy blogs and YouTube channels
Read about different treatment plans and seasonal practices
Beekeeping is a mix of science and intuition. You’ll make mistakes—everyone does. The important thing is to keep learning, stay curious, and support your bees the best you can.
Let me know if you’d like Chapter 14 too.
Here’s Chapter 14: Sustainable Beekeeping Practices :
Sustainable Beekeeping Practices
As you grow as a beekeeper, it’s worth thinking about sustainability—not just for your bees, but for your whole approach. Sustainable beekeeping means working with nature, not constantly fighting it.
Don’t Overharvest
Always leave plenty of honey for the bees. It’s their winter food. Taking too much puts your colony at risk and leads to unnecessary feeding or losses.
Let Bees Reproduce Naturally
Swarming is how bees reproduce as a colony. Instead of trying to prevent every swarm, plan for them. Make splits ahead of time or be ready to catch your own swarms. It's a natural way to expand and maintain strong genetics.
Raise Your Own Queens
Instead of buying queens every year, learn to raise a few from your own best colonies. Local queens tend to survive better and are already adapted to your climate and foraging conditions.
Use Natural Comb When Possible
Letting bees draw their own comb—especially in brood boxes—can reduce chemical buildup and give the bees more control over how they manage space. It's a bit messier, but often healthier.
Rethink Treatments
You don’t have to go treatment-free, but you should monitor closely. Use sugar rolls or alcohol washes to keep tabs on mites. Treat only when needed, and use methods that are effective but gentle (like formic or oxalic acid). Stay flexible and observant.
Plant for Pollinators
What grows around your bees matters. Plant native flowers, avoid pesticides, and leave some wild areas. The more bloom diversity and forage you offer, the healthier your bees (and local pollinators) will be.
Sustainability doesn’t mean doing it perfectly. It means doing it mindfully—with the long-term health of your bees, your land, and your community in mind.
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